For Tanjirou Kamado, Zenitsu Agatsuma, and Inosuke Hashibira, accompanying Kyoujurou Rengoku offers a chance to see an exemplary demon slayer in action. The Flame Pillar has taken on a mission aboard a train plagued by mysterious disappearances, after repeated efforts by the Demon Slayer Corps have failed to stop them. His task is to eliminate the threat before it claims more lives.
But the demons behind the disappearances have already set their plan in motion, without the slayers' knowledge. With two hundred passengers to protect, Rengoku and the young recruits must fight through the demonic threat with both their swords and their willpower. As the situation grows increasingly desperate, Tanjirou faces a test of his determination and commitment to duty.